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Estimate "between-axes" vs "within-axes heterogeneity of multivariate matrices

This time with a more-R oriented question:

Is the mrpp {vegan} package [1] useful in trying to check, or get a clue about 
the differences between- and within-axes (or variables or dimensions or 
columns) of a multivariate matrix?


The description explains:

" ...(MRPP) provides a test of whether there is a significant difference 
between two "or more groups of sampling units. ..."

"... difference may be one of location (differences in mean) or one of spread 
(differences in within-group distance) ..."

and 

"... Function mrpp operates on a data.frame matrix where rows are observations 
and responses data matrix. The response(s) may be uni- or multivariate. ..."


Question: what about the observations being actually the columns? Is a simple 
transposing of the matrix enough? Any other alternatives or hints?

Thak you, Nikos
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[1] <http://cc.oulu.fi/~jarioksa/softhelp/vegan/html/mrpp.html>


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Nikos: