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embedding expression into title in R plot

You instead need the `bquote` function. The `paste` function will only confuse things. In this particular instance it is embedding the literal as.character result of 'x_label'-value in a character object rather than in an expression-object. Since you have not offered a full example it remains unclear whether you want the words: "species" or "N1" rather than the values of those names.
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David Winsemius
Alameda, CA, USA