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lmer - error message

I think my problem is that I can't
Yes. Consider the collinearity between Age and Year, i.e. for a given
cohort (mos or all captured by "fishID" ?) they are essentially the
same variable with different units. So I would suspect the problem is
you are over fitting those.

A few suggestions:
Add one more to those: make sure your random effects are indeed
crossed. If they are nested (without knowing anything about your data,
just given their names that's a possibility), you could try nlme::lme.

Elai