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[R-meta] Model with intercept gives 0 heterogeneity but without intercept is ok

This doesn't sound right. Removing the intercept in this model says that the average effect must be 0 when X = 0. One can still estimates the variance components whether there is an intercept or not. They have different interpretations though, since the variances are estimated as deviations from a line that has an intercept of 0 or not.
Again, both variance components can be estimated. It just happens to be the case that sigma^2.1 is estimated to be essentially 0 in Model1.
Again, all variance components can be estimated whether the fixed part includes an intercept or not.
In this case, the model is identical whether you use '0 + X + gender' or 'X + gender', just the parameterization of the fixed part is different. Please see the link I posted.